Found in 10 samples

Bulgaria: Gorna Sushitsa (Late Miocene), Strumyani-2 (Late Miocene)

Greece: Pyrgos (Late Miocene)

Turkey: Akkasdagi (Late Miocene), Corakyerler (Late Miocene), Kemiklitepe (KTA) (Late Miocene), Kemiklitepe (KTB) (Late Miocene), Kemiklitepe (KTD) (Late Miocene), Esendere (Late Miocene), Kayaca (Late Miocene)

See also Miodiceros