Latitudinal range: 10.7° to 53.0°

Habitats: temperate broadleaf/mixed forest (6), tropical/subtropical dry broadleaf forest (8), tropical/subtropical moist broadleaf forest (1), temperate coniferous forest (1)

Found in 41 samples

Afghanistan: Nuristan Province

Bhutan: Lamai Goempa Research Preserve

Cambodia: Chhep Wildlife Sanctuary, Boh Dambang (Late Pleistocene)

China: Wanglang National Nature Reserve, Shennongjia National Nature Reserve, Nangunhe Nature Reserve, Guanyinshan Nature Reserve, Tengchong Section, Foping National Nature Reserve (2020), Fanjingshan National Nature Reserve, Mocun Cave (Late Pleistocene), Tianluoshan (Holocene), Cunkong Cave (Holocene), Lower Pubu Cave (Late Pleistocene), Wuyun Cave (Middle Pleistocene), Mohui Cave (Early Pleistocene), Ganxian Cave (Middle Pleistocene), Bailongdong (Middle Pleistocene), Yanlidong (Late Stage) (Late Pleistocene), Yanlidong (Early Stage) (Middle Pleistocene)

India: Dachigam National Park

Japan: Torihama (Holocene)

Laos: Tam Ngu Hao 2 (Middle Pleistocene)

Myanmar: Hukaung Valley Wildlife Sanctuary (Outside the Core), Kweenkoh, Mae Nyaw Kee South

Russia: Geographical Society Cave (Late Pleistocene), Imanay Cave (Pleistocene Layer 2) (Late Pleistocene), Imanay Cave (Pleistocene Layer 4) (Late Pleistocene)

Thailand: Khao Yai National Park, Khao Ang Rue Nai Wildlife Sanctuary, Cave of the Monk (Area 1) (Late Pleistocene), Ban Rai rockshelter (Holocene), Tham Lod rockshelter (Unit 1) (Late Pleistocene), Tham Lod rockshelter (Unit 2) (Late Pleistocene), Tham Lod rockshelter (Unit 3) (Late Pleistocene)

Ukraine: Medzhibozh 1 (horizon 16a) (Middle Pleistocene)

Vietnam: Duoi U'Oi (Late Pleistocene), Lang Trang (Cave II, breccia 5) (Late Pleistocene), Coc Muoi (Middle Pleistocene)

See also Ursus, Ursus americanus, Ursus arctos, Ursus bibersoni, Ursus boeckhi, Ursus deningeri, Ursus dolinensis, Ursus etruscus, Ursus kudarensis, Ursus maritimus, Ursus minimus, Ursus sp., Ursus spelaeus