PaleoDB taxon number: 286956
Full reference: R. W. Gess. 2013. The earliest record of terrestrial animals in Gondwana: a scorpion from the Famennian (Late Devonian) Witpoort Formation of South Africa. African Invertebrates 54:373-379
Sister taxa: none
Type specimen: AM7501, a claw. Its type locality is Waterloo Farm, which is in a Famennian paralic horizon in the Witpoort Formation of South Africa.